Zael is able to do several side quests as well as progress through the main story. They don't affect the story completion, but they do enable you to get more items, collectibles, or even abilities. Quests can only be found by talking to various different people, but other NPCs may mention that someone is looking for help. Quests can involve buying/finding items for someone, talking to a certain NPC, or something else.
There are also a notable amount of Easter Eggs and Tidbits that don't give awards besides some dialogue or backstory of Last Story and it's characters.
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